Peer-reviewed Publications


Walsh, J., I.J. Lovette, V. Winder, C.S. Elphick, B.J. Olsen, W.G. Shriver, and A.I. Kovach. 2017. Subspecies delineation amid phenotypic, geographic, and genetic discordance in a songbird. Molecular Ecology. 26(5):1242-1255.

Walsh, J. W.G. Shriver, M.D. Correll, B.J. Olsen, C.S. Elphick, T.P. Hodgman, R.J. Rowe, K.M. O’Brien, and A.I. Kovach. 2017. Temporal shifts in the saltmarsh-Nelson’s sparrow hybrid zone revealed by replicated demographic and genetic surveys. Conservation Genetics. 18(2):453-466.

Field, C.R., K.J. Ruskin, B. Benvenuti, A.C. Borowske, J.B. Cohen, L. Garey, T.P. Hodgman, R.A. Kern, E. King, A.R. Kocek, A.I. Kovach, K.M. O’Brien, B.J. Olsen, N. Pau, S.G. Roberts, E. Shelly, W.G. Shriver, J. Walsh, C.S. Elphick. 2017. Quantifying the importance of geographic replication and representativeness when estimating demographic rates, using a coastal species as a case study. Ecography.

Ruskin, K.J., M.A. Etterson, T.P. Hodgman, A.C. Borowske, J.B. Cohen, C.S. Elphick, C.R. Field, R.A. Kern, E. King, A.R. Kocek, A.I. Kovach, K.M. O’Brien, N. Pau, W.G. Shriver, J. Walsh, and B.J. Olsen. 2017. Seasonal fecundity is not related to geographic position across a species’ global range despite a central peak in abundance. Oecologia 183(1):291-301.


Walsh, J., B.J. Olsen, K.J. Ruskin, W.G. Shriver, K.M. O’Brien, and A.I. Kovach. 2016. Extrinsic and Intrinsic Factors Influence Fitness in an Avian Hybrid Zone. Biological Journal of the Linnean Society.

Walsh, J., W.G Shriver, B.J. Olsen, and A.I. Kovach. 2016. Differential Introgression and the Maintenance of Species Boundaries in an Advanced Generation Avian Hybrid Zone. BMC Evolutionary Biology. 16:65.


Walsh, J., J. Rowe, B.J. Olsen, W.G. Shriver, and A.I. Kovach. 2015. Genotype-Environment Associations Support a Mosaic Hybrid Zone Between Two Tidal Marsh Birds. Ecology & Evolution. 6(1):279-294.

Walsh, J., W.G. Shriver, B.J. Olsen, K.M. O’Brien, and A.I. Kovach. 2015. Relationship of phenotypic variation and genetic admixture in the Saltmarsh-Nelson’s sparrow hybrid zone. The Auk. 132(3): 704-716.

Kovach, A.I., J. Walsh, J. Ramsdell, and W.K. Thomas. 2015. Development of diagnostic microsatellite markers from whole-genome sequences of Ammodramus sparrows for assessing admixture in a hybrid zone. Ecology and Evolution. 5(11): 2267-2283.

Berlinsky, D.L., M.A. DiMaggio, T.S. Breton, J. Walsh, and A.I. Kovach. 2015. Peritoneal pigmentation in purebred and hybrid River Herring. Transactions of the American Fisheries Society. 144: 717-723.


Hill, J.M., J. Walsh, A.I. Kovach, and C. S. Elphick. 2013. Male-skewed nestling sex ratio in Saltmarsh Sparrows. Condor. 115(2): 411-420.


Walsh, J., A. I. Kovach, K. J. Babbitt, and K.M. O’Brien. 2012. Fine-scale population structure and asymmetrical dispersal in an obligate salt marsh passerine, the Saltmarsh Sparrow (Ammodramus caudacutus). The Auk. 129 (2): 247-258.


Walsh, J., A.I. Kovach, O.P. Lane, K.M. O’Brien, and K.J. Babbitt. 2011. Genetic barcode RFLP analysis of the Nelson’s and Saltmarsh Sparrow Hybrid Zone.  Wilson Journal of Ornithology. 123(2): 316-322.

Popular Press Articles

Kovach, A.I. and J. Walsh. 2012. Interactions of two salt marsh-dwelling sparrows in New England: past, present and future. Bird Observer. 40 (1): 25-32.